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Williams was given the room to ad-lib (at some points in the script, they didn't even bother to write dialogue for him) and Genie lead animator Eric Goldberg got to go through all of Williams' ad-libs, pick the best ones, have his team animate to those, and assemble them into what we eventually saw on film.
In short cartoons, Jack Mercer in "Popeye the Sailor Meets Ali-Baba's Forty Thieves" (1937) is probably my nomination. Until about 1940 or so, Popeye cartoons - unlike most American sound cartoons - were dubbed entirely in post, after everything was finished. As such, Jack Mercer got to do those infamous "under-the-breath mutterings" that Popeye is known for as commentary o nthe finished film, which he is watching while doing the dialogue. "Popeye Meets Ali Baba" is probably his tour-fe-force - "This writin's wrotten rotten if ya happen to ask me, huh?" https://www.youtube.com/watch?v=miw_Jw7j2sE
A list of others in film, short cartoons and TV that immediately come to mind: Mel Blanc in basically anything is an obvious one, usually in Chuck Jones-directed Looney Tunes and Merrie Melodies like the "Duck, Rabbit, Duck!" (1952) - "pronoun trouble" Rob Paulsen, Trish MacNell, Jess Harnell, and Bernadette Peters in "Animaniacs" Gary Anthony Williams in "The Boondocks" Paul Lynde in "Charlotte's Web" ("It say-yas...'crun-chee'") Casey Kasem on "Scooby-Doo, Where are You!" Frank Welker and Grey DeLisle on "Scooby-Doo! Mystery Incorporated"
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